Career & Publications

Career Path

Current position:

2024-present NERC Independent Research Fellow, University of Exeter

Past positions:

2021-2024 Herchel Smith Postdoctoral Fellow, Department of Zoology, University of Cambridge

2021-2024 Homerton College Research Associate, University of Cambridge

2019-2020 Biodiversity Research Fellow, Museum of Comparative Zoology & OEB, Harvard University


2015-2019 DPhil Zoology, University of Oxford

Included 1 year as a visiting researcher (ISTE, University of Lausanne) 2017-2018

2010-2014 MSci Earth Sciences, University of Cambridge


36. Wu, Y.*, Pates, S.*, Pauly, D., Zhang, X., Fu, D. 2023. Rapid growth in a large Cambrian apex predator. National Science Review. 0: nwad284. *joint first authors

35. Pates, S.* and Zamora, S.* 2023. Large euarthropod carapaces from a high latitude Cambrian (Drumian) deposit in Spain. Royal Society Open Science. 10:230935. *joint first authors

34. Kimmig, J., Pates, S., LaVine, R., Krumenacker, L. J., Whitaker, A. F., Strotz, L. C., Jamison, P. G., Gunther, V. G., Gunther, G., Witte, M., Daley, A. C. and Lieberman, B. S. 2023. New soft-bodied panarthropods from diverse Spence Shale (Cambrian; Miaolingian; Wuliuan) depositional environments. Journal of Paleontology. 

33. Ma, J., Pates, S., Wu, Y., Lin, W., Liu, C., Wu, Y., Zhang, M. and Fu, D. 2023. Ontogeny and brooding strategy of the early Cambrian arthropod Isoxys minor from the Qingjiang biota. Frontiers in Ecology and Evolution. 11:1174564. 

32. Botting J.P., Muir, L.A., Pates, S., McCobb, L.M.E., Wallet, E., Willman, S., Zhang, Y. and Ma, J. 2023. A Middle Ordovician Burgess Shale-type fauna from Castle Bank, Wales (UK). Nature Ecology & Evolution. 7. 666-674. 


31. Pates, S., Botting, J.P., Muir, L.A. and Wolfe, J.M. 2022. Ordovician opabiniid-like animals and the role of the proboscis in euarthropod head evolution. Nature Communications, 13: 6969.

30. Wu, Y., Pates, S., Ma, J. Lin, W. Wu, Y., Fu, D. and Zhang, X. 2022. Addressing the Chengjiang Conundrum: a palaeoecological view on the rarity of early Cambrian hurdiid radiodonts in the most diverse early Cambrian Lagerstätte. Geoscience Frontiers, 13: 101430.

20. Bicknell, R. D. C., Holmes, J. D., Pates, S., Garcia-Bellido, D. C. and Paterson, J. 2022. Cambrian carnage: trilobite predator-prey interactions in the Emu Bay Shale of South Australia. Palaegeography, Palaeoclimatology, Palaeoecology. 591: 110877.

28. Pates, S.*, Wolfe, J. M.*, Lerosey-Aubril, R., Daley, A. C. and Ortega- Hernández, J. 2022 New opabiniid diversifies the weirdest wonders of the euarthropod stem group. Proceedings of the Royal Society B, 289: 20212093. *joint first authors


27. Jiao, D.G.*, Pates, S*., Lerosey-Aubril, R., Ortega-Hernández, J., Yang, J., Lan, T. and Zhang, X.G. 2021. New multipodomerous appendages of stem group euarthropods from the Cambrian (Stage 4) Guanshan Konservat-Lagerstatte. Royal Society Open Science, 8: 211134. *joint first authors

26. Pates, S., Daley, A. C., Legg, D. A. and Rahman, I. A. 2021. Vertically migrating Isoxys and the early Cambrian biological pump. Proceedings of the Royal Society B, 288: 20210464.

25. Pates, S., Daley, A. C., Edgecombe, G. D., Cong, P., and Lieberman, B. S. 2021. Systematics, preservation, and biogeography of radiodonts from the southern Great Basin, USA, during the upper Dyeran (Cambrian Series 2, Stage 4). Papers in Palaeontology. 7, 235-262.

24. Pates, S.*, Lerosey-Aubril, R.*, Daley, A. C., Kier, C., Bonino, E. and Ortega-Hernández. 2021. The diverse radiodont fauna from the Marjum Formation of Utah, USA (Cambrian: Drumian). PeerJ. e10509. *joint first authors

23. Jiao, D.G., Pates, S., Lerosey-Aubril, R., Ortega-Hernández, J., Yang, J., Lan, T. and Zhang, X.G., 2021. The endemic radiodonts of the Cambrian Stage 4 Guanshan Biota of South China. Acta Palaeontologica Polonica, 66, 255-274.


22. Pates S., Botting J. P., McCobb L. M. E., Muir L. A. 2020 A miniature Ordovician hurdiid from Wales demonstrates the adaptability of Radiodonta. Royal Society Open Science.7: 200459.

21. Bicknell, R. D. C., and Pates, S. 2020. Pictoral atlas of fossil and extant horseshoe crabs with focus on Xiphosurida. Frontiers in Zoology. 8, 98.

20. Bicknell, R. D. C., and Pates, S. 2020. Exploring abnormal Cambrian-aged trilobites in the Smithsonian collection. PeerJ. 8, e8453.

19. Lerosey-Aubril, R., Kimmig, J., Pates, S., Skabelund, J., Weug, A., and Ortega- Hernández, J. 2020. New exceptionally-preserved arthropods from the Drumian Wheeler Formation Konservat-Lagerstatte of the House Range of Utah. Papers in Palaeontology. 6, 501-531.


18. Ortega- Hernández, J., Lerosey-Aubril, R., and Pates, S. 2019. Proclivity of nervous system preservation in Cambrian Burgess Shale-type deposits. Proceedings of the Royal Society B. 286, 20192370.

17. Bicknell, R. D. C., and Pates, S. 2019. Xiphosurid from the Tournaisian (Carboniferous) of Scotland confirms deep origin of Limuloidea. Scientific Reports. 9, 17102.

16. Pates, S., Daley, A. C., and Butterfield, N. J. 2019. First report of paired ventral endites in a hurdiid radiodont. Zoological Letters. 5, 18.

15. Pates, S. and Bicknell, R. D. C. 2019. Elongated thoracic spines as potential predatory deterrents in olenelline trilobites from the lower Cambrian of Nevada. Palaeogeography, Palaeoclimatology, Palaeoecology. 516, 295-306.

14. Bicknell, R. D. C. and Pates, S. 2019. Abnormal extant xiphosurids in the Yale Peabody Museum Invertebrate Zoology collection. Bulletin of the Yale Peabody Museum of Natural History. 60, 41-53.

13. Pates, S. and Daley, A. C. 2019. The Kinzers Formation (Pennsylvania, USA): the most diverse assemblage of Cambrian Stage 4 radiodonts. Geological Magazine. 156, 1233-1246.

12. Guo, J., Pates, S.*, Cong, P., Daley, A. C., Edgecombe, G. D., Chen, T. and Hou, X. G. 2019. A new radiodont (stem Euarthropoda) frontal appendage with a mosaic of characters from the Cambrian (Series 2 Stage 3) Chengjiang biota. Papers in Palaeontology. 5, 99–110. *corresponding author

11. Bicknell, R. D. C., Pates, S., and Botton, M. L. 2019. Euproops danae cluster confirms deep origin of gregarious behavior in xiphosurids. Arthropoda Selecta. 28, 549-555.

10. Lerosey-Aubril, R. and Pates, S. 2018. New suspension-feeding radiodont suggests evolution of microplanktivory in Cambrian macronekton. Nature Communications. 9, 3774.


9. Pates, S., Daley, A. C., and Lieberman, B. S. 2018. Hurdiid radiodontans from the middle Cambrian (Series 3) of Utah. Journal of Paleontology. 92, 99-113.

8. Daley, A. C., Antcliffe, J. B., Drage, H. B. and Pates, S. 2018. Early fossil record of Euarthropoda and the Cambrian Explosion. PNAS. 115, 5323 – 5331.

7. Bicknell, R. D. C., Pates, S. and Botton, M. L. 2018. Abnormal xiphosurids, with possible application to Cambrian trilobites. Palaeontologia Electronica. 21, 1–17.

6. Cong, P., Edgecombe, G. D., Daley, A. C., Guo, J., Pates, S., and Hou, X. G. 2018. New radiodonts with gnathobase-like structures from the Cambrian Chengjiang Biota and implications for the systematics of Radiodonta. Papers in Palaeontology. 4, 605-621.

5. Pates, S., Daley, A. C., and Ortega-Hernández, J. 2018. Response to Comment on “Aysheaia prolata from the Wheeler Formation (Cambrian, Drumian) is a frontal appendage of the radiodontan Stanleycaris” with the formal description of Stanleycaris. Acta Palaeontologica Polonica. 63, 105-110.


4. Pates, S., Bicknell, R.D.C., Daley, A.C., and Zamora, S. 2017. Quantitative analysis of repaired and unrepaired damage to trilobites from the Cambrian (Stage 4, Drumian) Iberian Chains, NE Spain. PALAIOS. 32, 750-761.

3. Pates, S., Daley, A. C., and Ortega-Hernández, J. 2017. Aysheaia prolata from the Wheeler Formation (Cambrian, Drumian) is a frontal appendage of the radiodontan Stanleycaris. Acta Palaeontologica Polonica. 62, 619-625.

2. Pates, S., and Daley, A. C. 2017. Caryosyntrips: a radiodontan from the Cambrian of Spain, USA and Canada. Papers in Palaeontology. 3, 461-470.

1. Legg, D., and Pates, S. 2017. A restudy of Utahcaris orion (Euarthropoda) from the Spence Shale (Middle Cambrian, Utah, USA). Geological Magazine. 154, 181-186.

Book chapter

1. Bicknell, R. D. C., Pates, S, Kaiser, D., Zakrzeqski, S. and Botton, M. L. 2022. Applying records of extant and extinct horseshoe crab abnormalities to xiphosurid conservation. In: Tanacredi J. T. et al. International Horseshoe Crab Conservation and Research Efforts: 2007-2020. Springer International Publishing, pp 85-104.

Field guide

1. Zamora, S., Alvaro, J. J., Esteve, J., and Pates, S. 2019. Day 2 December 19th, Cambrian invertebrate faunas from the Badules Unit: Purujosa and Murero localities. Palaeontological Association Pre-Conference Field Trip: Palaeozoic fossil assemblages from the Iberian Chains. Eds. Zamora, S. and Rabano, I. Cuadernos del Museo Geominero, 31. Instituto Geológico y Minero de España, Madrid. ISBN: 978-84-9138-095-5